A girl named Sakura flees after her fox companion, Shizuku, is captured. After escaping herself, she ends up in the human world, where Ichika Usami finds her. Sakura explains she comes from "Sakuragahara," a world inspired by ancient Japan now under threat from a sinister figure called Karasu Tengu, who has stolen everything beautiful from her home. She is searching for the Pretty Cure, who embody the concepts of Sweet, Jewels, and Keys. Ichika reveals she knows the KiraKira PreCure team, representing sweets. When two of Karasu Tengu’s minions arrive in the human world to track Sakura down, the KiraKira team is ambushed-only to be rescued by the Maho Girls PreCure, representing jewels, and the Go! Princess PreCure, representing keys. Once united with Sakura, the three teams must venture into Sakuragahara to save Shizuku and free her world from Karasu Tengu’s grip.

Pretty Cure Dream Stars!
A girl named Sakura flees after her fox companion, Shizuku, is captured. After escaping herself, she ends up in the human world, where Ichika Usami finds her. Sakura explains she comes from "Sakuragahara," a world inspired by ancient Japan now under threat from a sinister figure called Karasu Tengu, who has stolen everything beautiful from her home. She is searching for the Pretty Cure, who embody the concepts of Sweet, Jewels, and Keys. Ichika reveals she knows the KiraKira PreCure team, representing sweets. When two of Karasu Tengu’s minions arrive in the human world to track Sakura down, the KiraKira team is ambushed-only to be rescued by the Maho Girls PreCure, representing jewels, and the Go! Princess PreCure, representing keys. Once united with Sakura, the three teams must venture into Sakuragahara to save Shizuku and free her world from Karasu Tengu’s grip.
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